Fixed-term

We are offering a PhD fellowship in cancer immunology and targeted cancer therapy commencing 31 August 2026. Our group and research Research at the Centre for translational Medicine and Parasitology (CMP) is focused on malaria and recently on using the learnings from malaria during pregnancy to develop novel cancer therapies. All activities are organized in thematically defined teams, each led by a senior scientist. The team leaders, the Head of CMP, and a technician representative form the daily leadership of CMP. CMP is part of a well-established international scientific network, composed of scientists in Europe, Africa, America, and Australia. More than 70 scientists and technicians are affiliated at CMP. Our research team spans development of vaccines against malaria and other infectious diseases as well as research and development of novel cancer therapies and diagnostic tools and basic research into how tumors avoid immune attack. Our research facilities are located at the Panum Institute in the Mærsk Tower. Project description The PhD project is focused on the basic role of tumor microenvironment in cancer and in particular on the role of onco-fetal chondroitin sulfates in tumor immune evasion. The project entails studies on human tissue and human cell lines, as well as advanced animal models including genetic knockout studies and GEMM models of cancer. Principal supervisor is Professor Ali Salanti, Department of Immunology and Microbiology, University of Copenhagen, e-mail: salanti@sund.ku.dk Start 31 August 2026 Duration 3 years as a PhD student Job description Your key tasks as a PhD student at SUND are: Carrying through an independent research project under supervision. Completing PhD courses or other equivalent education corresponding to approximately 30 ECTS points. Participating in active research environments including a stay at another research team. Obtaining experience with teaching or other types of dissemination related to your PhD project Teaching and disseminating your knowledge. Writing a PhD thesis on the grounds of your project PhD Programme Description The PhD programme is a three-year full-time research education within the framework of the 5+3 scheme, intended for candidates holding a relevant academic degree within the project’s subject area (e.g., [relevant degrees]). Qualifications If you have completed your academic education in Denmark, the admission to the PhD programme requires a relevant Master’s degree (equivalent to 180 ECTS BSc + 120 ECTS MSc). If you have an academic education from abroad, you must have either a degree equivalent to a Danish master’s degree (180 ETCS + 120 ECTS MSc), or at a minimum, a bachelor’s degree equivalent to a Danish bachelor’s degree (180 ECTS) and be academically qualified at master’s level. As part of the assessment of whether you meet the educational requirements, you must also achieve a positive academic evaluation and be assessed as the best qualified for the position. Other important criteria are: The grade point average achieved Professional qualifications relevant to the PhD project Previous publications Relevant work experience Other professional activities A curious mind-set with a strong interest in oncology Language skills Place of employment The place of employment is at the Department of Immunology and Microbiology, Faculty of Health and Medical Sciences, University of Copenhagen. We offer creative and stimulating working conditions in dynamic and international research environment. Our research facilities include modern laboratories and access to state-of-the-art core facilities. Terms of employment The average weekly working hours are 37 hours per week. The position is a fixed-term position limited to a period of 3 years. The start date is 31 August 2026 The employment is conditioned upon the applicant’s success­ful enrolment as a PhD student at the Graduate School at the Faculty of Health and Medical Sciences, University of Copenhagen. This requires submission and acceptance of an application for the specific project formulated by the applicant. The PhD study must be completed in accordance with The Ministerial Order on the PhD programme (2025) and the Faculty’s rules on achieving the degree. Salary, pension and terms of employment are in accordance with the agree­ment between the Ministry of Taxation and The Danish Confederation of Professional Associations on Academics in the State. Depending on seniority, the monthly salary starts at approximately 31,242 DKK/Roughly 4,180 EUR (November 2025 level) plus pension. Questions For specific information about the PhD fellowship, please contact the principal supervisor. General information about PhD studies at the Faculty of Health and Medical Sciences is available at the Graduate School’s website: https://healthsciences.ku.dk/phd/guidelines/ Application procedure Your application must be submitted electronically by clicking ‘Apply now’ below. The application must include the following documents in PDF format: Motivated letter of application (max. one page) CV incl. education, experience, language skills and other skills relevant for the position Certified copy of original Master of Science diploma and transcript of records in the original language, including an authorized English translation if issued in other language than English or Danish. If not completed, a certified/signed copy of a recent transcript of records or a written statement from the institution or supervisor is accepted. As a prerequisite for a PhD fellowship employment, your master’s degree must be equivalent to a Danish master’s degree. We encourage you to read more in the assessment database: https://ufm.dk/en/education/recognition-and-transparency/find-assessments/assessment-database. Please note that we might ask you to obtain an assessment of your education performed by the Ministry of Higher Education and Science Publication list (if possible) Application deadline is 7 May 2026, 23.59 CET. We reserve the right not to consider material received after the deadline, and not to consider applications that do not live up to the abovementioned requirements. The further process After the expiry of the deadline for applications, the authorized recruitment manager selects applicants for assessment on the advice of the hiring committee. All applicants are then immediately notified whether their application has been passed for assessment by an unbiased assessor. The assessor makes a non-prioritized assessment of the academic qualifications and experience with respect to the above-mentioned area of research, techniques, skills and other requirements listed in the advertisement. Once the assessment work has been completed, each applicant has the opportunity to comment on the part of the assessment that relates to the

PhD position There is a vacancy for up to two PhD Research Fellows at the Michael Sars Centre affiliated with the project “Molecular, cellular and developmental marine living systems in a variable ocean”. The position is for a fixed-term period of 4 years, with 25 percent of the time each year dedicated to required career-promoting work. The position is financed by the University of Bergen. About the project/work tasks: The Christiaen lab uses a wide range of approaches, including molecular, cellular and embryological techniques, genomics including from single cells, genome engineering and computational biology methods to study development and regeneration, at a systems level, in tunicates. Tunicates, especially the genus Ciona, offer unique opportunities to leverage cutting edge experimental and computational techniques and approaches to study chordate development with a cellular resolution and at the genome scale. Specifically, efforts using systematic CRISPR/Cas9-mediated mutagenesis, high-content imaging, data science and computational modelling are used to study the biomolecular networks governing cell fate specification and cellular behaviours, such as collective migration and oriented asymmetric cell divisions. New directions will leverage controlled culture conditions, the natural diversity of tunicate populations, single cell genomics, quantitative imaging, CRISPR/Cas9, and data science to probe the impact of environmental challenges on developing embryos at the systems scale. The projects offer opportunities for collaborations both locally and internationally. Qualifications and personal qualities: Applicants must hold a master’s degree or equivalent education in biology or must have submitted his/her master’s thesis for assessment prior to the application deadline. It is a condition of employment that the master’s degree has been awarded. Experience in either molecular cloning, developmental and cell biology, quantitative image analysis, and/or genomics and computational biology is required. Experience with ‘emerging’ model organisms in the fields of genetics, developmental biology, cell biology and/or molecular physiology is an advantage. Experience in analysing next-generation sequencing datasets is an advantage. We are seeking a candidate with interests in developmental, cell and/or systems aspects of animal development. Impeccable work ethics, rigor and discipline to complete defined tasks, including detailed reporting, are required. Background knowledge of the relevant literature is an advantage. Creative thinking and the ability to identify important questions and design scientific approaches is an advantage. Organizational and communication skills are advantages. A strong motivation to perform research at an internationally competitive level is expected. Applicants must be able to work independently and in a structured manner and demonstrate good collaborative skills. Applicants must have good written and oral English skills. Personal and relational qualities will be emphasized. Research experience, ambitions and potential will also count when evaluating the candidates. Special requirements for the position The University of Bergen is subjected to the regulation for export control system. The regulation will be applied in the processing of the applications. About the PhD position: The duration of the PhD Research Fellowship is 4 years, with 25 percent of the time each year dedicated to required career-promoting work related to research, teaching, and dissemination of results. The employment period may be reduced if you have previously been employed in a qualifying post (e.g. research fellow, research assistant). About the research training: As a PhD Research Fellow, you must participate in an approved educational programme for a PhD degree within a period of 3 years. The deadline for applying for admission to the PhD programme at The Faculty of Science and Technology is 2 months after you start your position or after the start of the research project that will lead to the PhD degree. It is a condition that you satisfy the enrolment requirements for the PhD programme at the University of Bergen. We can offer: Exciting development opportunities as part of your role in a strong professional environment Salary as a PhD research fellow (code 1017) in the state salary scale. This currently amounts to an annual salary of NOK 568 700 before taxes. Further salary increases are made according to the length of service in the position. Enrolment in the Norwegian Public Service Pension Fund Good welfare benefits Your application must include: a brief account of the applicant’s research interests and motivation for applying for the position a detailed justification of your qualifications in relation to the listed requirements, explaining how you meet each criterion. Additionally, you should address the “advantages” mentioned above, as these will be taken into account when ranking the candidates. the names and contact information for two referees. One of these should be the main advisor for the master’s thesis or equivalent thesis CV transcripts and diplomas showing completion of the bachelor’s and master’s degrees. If you have not yet completed your master’s degree, please submit a statement from your institution confirming that the master’s thesis has been submitted relevant certificates/references approved documentation of proficiency in English (if required, cf. English language requirements for PhD admission) a list of any works of a scientific nature (publication list with links) The application and appendices with certified translations into English or a Scandinavian language must be uploaded at Jobbnorge. General information: For further details about the position, please contact Group Leader Lionel Christiaen, email: lionel.christiaen@uib.no or Head of Administration Kristina Håvås Hanson, email: kristina.hanson@uib.no. For HR related questions contact HR adviser Veronica Stålhand, Phone: +47 55 58 43 74, email: veronica.stalhand@uib.no. Diversity is a strength that enables us to solve our tasks even better. UiB therefore needs qualified employees regardless of gender, ethnicity, religion, worldview, disability, sexual orientation, gender identity, gender expression, and age. We encourage women to apply. If multiple applicants have approximately equivalent qualifications, the rules pertaining to moderate gender quotas shall apply. The University of Bergen applies the principle of public access to information when recruiting staff for academic positions. Information about applicants may be made public even if the applicant has asked not to be named on the list of persons who have applied. The applicant must be notified if the request to be omitted is not met. The successful applicant must comply with the guidelines that apply to the position at all times. We encourage applicants with disabilities, immigrant backgrounds, or gaps in their CV to apply. By indicating such circumstances in your application, you may receive favourable consideration. We ensure that at

The position The Faculty of Biosciences, Fisheries and Economics (BFE), UiT-The Arctic University of Norway in close collaboration with SINTEF Ocean (Tromsø, Trondheim) and SINTEF Industry (Oslo) has a PhD position vacant in application of biodegradable materials in marine aquaculture – for applicants wishing to obtain the degree of Philosophiae Doctor (PhD). The position is attached to Centre for Research-based Innovation Dsolve at the Norwegian College of Fishery Science (NCFS). The position is for a period of four years. The nominal length of the PhD programme is three years. The fourth year is distributed as 25 % each year and will consist of teaching and other duties. The objective of the position is to complete research training to the level of a doctoral degree. Admission to the PhD programme is a prerequisite for employment, and the programme period starts on commencement of the position. The Norwegian College of Fishery Science (NCFS) undertakes research and research-based teaching of high national and international quality in an interdisciplinarty environment. Research activities are provisionally organized into groups of Living Marine Resources, Food science, Fish Health, Marine Drug Discovery, Marine Resource Management and Development, Environmental and Resource Economics. The position is within the center for research-based innovation: SFI Dsolve – Biodegradable plastics for marine applications which includes SINTEF Ocean and SINTEF Industry as well as several other research and industry partners. The candidate shall collaborate with industrial partners within marine aquaculture in the SFI. Supervisors for the candidate will be from UiT-Faculty of Biosciences, Fisheries and Economy, SINTEF Ocean, SINTEF Industry, and other partners. The workplace is at UiT in Tromsø, although the candidate is expected to spend a proportion of the time partly in Trondheim at SINTEF Ocean and potentially in corporation with SINTEF Industry in Oslo. SINTEF Ocean and SINTEF Industry are in possession of facilities for weathering experiments and mechanical tests, including labs for the investigation of biodegradation in natural sea water and marine sediments, polymers and composites research. The applicant is expected to work in co-operation with aquaculture companies in the Dsolve consortium in order to gather data for further analyses. The elected candidate must be able to start in the position within a reasonable time after receiving the offer. The project / The position’s field of research The PhD project is within the SFI Dsolve financed by the Research Council of Norway. This is a multi-disciplinary project concentrated around the development and implementation of biodegradable plastics in fisheries and aquaculture. Norway’s salmon farming industry is a major export sector, yet the issue of microplastic pollution from aquaculture has received less attention compared to fisheries. However, this is now changing, and this industry is expected to implement measures to reduce microplastic emissions in the coming years. This PhD project will contribute to these efforts by investigating strategies to mitigate microplastic pollution from aquaculture, with a particular focus on single-use ropes and feeding tubes used in salmon farming. The PhD candidate will select a primary research focus within the overarching theme of microplastic reduction in aquaculture. The primary aspect must address the use of single-use ropes in aquaculture, exploring their usage patterns, methods to reduce their consumption, and possibilities for implementation of biodegradable alternatives. The project work will involve field experiments conducted at aquaculture facilities in collaboration with Dsolve industry partners, and laboratory research to assess material properties and degradation processes. Furthermore, interdisciplinary collaboration with experts in chemistry, physics, marine biology, aquaculture, fisheries technology, and industry stakeholders. The applicants must present a description outlining the academic basis of the PhD project. The project description shall be within 3-5 pages, literature references included. It must include a description of the topic, research question(s) and reasoning of the choices. It should also indicate the methodologies to be used. The final project description will be developed in cooperation with the supervision team. Qualifications Required qualifications: Master’s degree or foreign equivalent in a relevant field, such as aquaculture or fishery technology, environmental science, material science. Qualification with a master’s degree is required before commencement in the position. If you are near completion of your master’s degree, you may still apply and submit a draft version of the thesis and a statement from your supervisor or institution indicating when the degree will be obtained. You must still submit your transcript of grades for the master’s degree with your application. The applicant must prove some educational or work experience related to ocean aquaculture (fish farming) Documented knowledge in Norwegian or another Scandinavian language. The ability to speak Norwegian or another Scandinavian language is considered especially necessary during field work on ocean aquaculture installations. Documented fluency of English. Nordic applicants can document their English capabilities by attaching their high school diploma. Desired qualifications: Work experience from Nordic marine aquaculture (fish farming) industry will be considered a merit. Knowledge of the Nordic aquaculture industry, with special emphasis on salmon farming. Knowledge and experience in systematic testing in laboratory or in field work settings of materials used in aquaculture. Experience from or interest in sustainability, marine plastic littering, aquaculture technology, and interdisciplinary research. Industrial experience, e.g., production of gear components used in fisheries and/or aquaculture. It will be considered a merit if the applicant has safety course(s) from fisheries or aquaculture. Other qualifications The preferred candidate must have excellent analytical and collaboration skills as well as an ability to work both independently and in teams. Furthermore, it will be of advantage if the chosen candidate can prove experience from writing and publications in English. In the assessment, the emphasis is on the applicant’s potential to complete a research education based on the master’s thesis or equivalent, and any other scientific work. The project description will also be considered. In addition, professional experience and other experience of significance for the completion of the doctoral programme will be taken into consideration. We will also emphasize motivation and personal suitability for the position. We are looking for candidates who: Have good communication and interaction with colleagues and students Wants to contribute to a good working environment As many people as possible should have the opportunity to undertake organized research training. If you already hold a PhD or have equivalent competence, we will not appoint you to this position. Admission to the PhD programme For employment in the PhD position, you must be qualified for admission to the PhD programme at the Faculty of Biosciences, Fisheries and Economics and participate in organized doctoral studies within the employment period. Admission normally requires: A bachelor’s degree of 180 ECTS and a master’s degree of 120 ECTS, or an integrated master’s degree of 300 ECTS. A master’s thesis with a scope corresponding to at least 30 ECTS for a master’s degree of 120 ECTS. A master’s thesis with a scope corresponding to at least 20 ECTS for an integrated master’s degree of 300 ECTS. Grade requirements applies both to the master thesis as well as courses part of the master’s degree. The minimum requirements are grade B or better on the master’s degree, and an average grade of B or better on courses that are part of the master’s degree. For integrated master’s degrees, applicants are required to have achieved a grade of B or higher on their master’s thesis and an overall average grade of C or higher across all courses comprising the integrated degree programme. A more detailed description of admission requirements can be found here. If you are employed in the position, you will be provisionally admitted to the PhD programme. Application for final

About the position Faculty of Biosciences and Aquaculture, Nord University, Norway, invites applications from outstanding candidates for a three‑year PhD position in zooplankton molecular ecology. The successful candidate will be part of one of the ecology divisions, located in Bodø. Calanus species play a crucial role in coastal ecosystems in the North Atlantic. You will use historical DNA from the zooplankton collections at Nord University to investigate how environmental and climatic factors influence the distribution of Calanus species in local fjords. This project will contribute to a better understanding of the interactions between climate change and plankton communities, and improve forecasting of changes in food‑web structure in Norwegian fjords. We are looking for motivated, creative and ambitious applicants with an excellent academic record and the ability to collaborate with members of a multidisciplinary team. Qualification requirements The candidate should have an MSc degree (or equivalent) in biology, genetics, molecular ecology or equivalent fields. Grade average of B or better (or equivalent score) both for the programme in total and the master’s thesis. The grades at the qualifying MSc examination need to be B or better (ECTS scale from A-E) to be admitted to the PhD program. See admission requirements: PhD in Biosciences. Employment is conditional upon admission to the faculty’s doctoral programme. The final project outline must be approved no later than three months after the start of employment. Termination may be considered if admission to the doctoral program is not available within this deadline, due to insufficient progress or failure to meet required duties. For admission requirements and regulations, see our web page. Good written and oral English skills are required both written and oral. For non‑native speakers, proficiency in English is essential and must be documented by recognized certificates. Good knowledge and practical experience in molecular ecology, population genetics, and statistical analyses is required. Interest and prior practical experience with molecular ecology of zooplankton is preferred. Practical experience with Calanus and/or DNA extraction from historical collections is highly desirable. Candidates who already have a PhD degree will not be considered for the position. Qualified applicants will be ranked according to the following criteria: Theme and perspectives from the applicant’s master’s study The applicant’s description of motivation for the doctoral work and opportunities for completion Relevant experience from research work Ability to work independently, goal-oriented and systematically Personal suitability We offer The annual salary for doctoral researchers (code 1017) is NOK 550 800,-. Advantages within the Government Pension Fund for loans, insurance and pension Creative and collegial working environment A workplace with progressive development Flexible working hours Active university sports team General information The person who is appointed must abide by the applicable laws, agreements, and directives. The responsibilities and duties associated with the position may change with future reorganization at Nord University. Nord University is subject to the Security Act, which sets requirements for the handling of classified and sensitive information. To increase security in the hiring process, Nord may conduct background checks on relevant candidates. Please note that all our candidates may be asked questions necessary in this context. This includes questions about any potential connection to countries that PST has identified as key threat actors in its annual threat assessment. The state workforce should, to the greatest extent possible, reflect the diversity of the population. Therefore, Nord University encourages qualified candidates with disabilities, gaps in their CV, immigrant backgrounds, and other diverse life experiences to apply. Information about disabilities or gaps in the CV may be used for anonymized registration purposes. For this position, Nord University is cautious about hiring individuals with connections to  countries that PST has identified as key threat actors in its annual threat assessment. Application process Applicants will be evaluated by an internal expert committee. Relevant applicants will be invited for an interview, trial lecture or other presentations. For more information, see Nord`s website . The Faculty reserve the right not to hire applicants who may be scientifically qualified but who do not meet other criteria such as language, educational background or personal characteristics Application The application should include: Application letter with description of your motivation for applying for the position Complete CV (education, work and teaching experience and overview of any scientific publications) Copy of diploma for both bachelor and master education and relevant certificates Applicants with education from countries outside Scandinavia with diplomas issued in a language other than English, must enclose witnessed diplomas in both the original language and authorized English translations Applicants from outside the EU/EEA must submit HK-dir approval of their master´s degree Copy of the applicant’s master’s thesis Name and contact information of 1-3 references (name, your relation to the person, email and phone number) Other activity that is considered relevant for the position, and wishes to be taken into account in the assessment All documents must be uploaded as attachments as a PDF files to the electronic application form. It is the applicant’s responsibility to submit the complete documentation within the deadline. After the application period has expired a list of applicants will be published. Note that, in special cases, the identity of particular applicants to a position may be made public even when that applicant has requested that their application be processed confidentially. Applicants requesting an exemption from publication of their identity must explain their reasons for doing so in their application. In this case, advance notice will be given in the event of a decision to make any information about the applicant public. Apply Now

About the PhD Fellowship Faculty of biosciences and aquaculture, Nord University, Norway, invites applications from outstanding candidates for a three-year PhD position in animal science. Join an innovative PhD project tackling one of dairy farming’s most debated welfare challenges: early cow– calf separation. By the development of AI-based tools to measure cow–calf interactions, the project aims to generate new scientific insights into CCC systems and contribute to future welfare policy for cow-calf housing. If you are interested in the cross-section of animal behaviour research and automatic animal monitoring – this project is for you! The project will include an animal trial at the Livestock Production Research Centre, NMBU, Ås, Norway and the successful candidates will be expected to travel for research stays, courses and conferences. The successful candidate will be part of one of the academic divisions Animal Science, Production and Welfare. This is a three-year contract based in Steinkjer, Trøndelag, Norway. Qualification requirements A completed master’s degree that meets the admission requirements, relevant academic backgrounds include ethology, animal or veterinary science, computer engineering or other closely related disciplines Good English skills, both written and oral Skills Experience in conducting farm animal trials and knowledge of fundamental ethology in farm animals Knowledge of data handling, with some exposure to big-data workflows or modelling Good communication and interpersonal skills Candidates who already have a PhD degree will not be considered for the position. Qualified applicants will be ranked according to the following criteria: Theme and perspectives from the applicant’s master’s project The applicant’s description of motivation for the doctoral work and opportunities for completion Personal suitability and scientific integrity Relevant experience from research work Ability to work independently, goal-oriented and systematically We offer The annual salary for doctoral researchers (code 1017) is NOK 550 800,-. Advantages within the Government Pension Fund for loans, insurance and pension Creative and collegial working environment A workplace with progressive development Flexible working hours Active university sports team General information The person who is appointed must abide by the applicable laws, agreements, and directives. The responsibilities and duties associated with the position may change with future reorganization at Nord University. Nord University is subject to the Security Act, which sets requirements for the handling of classified and sensitive information. To increase security in the hiring process, Nord may conduct background checks on relevant candidates. The public sector workforce is expected to reflect the diversity of the population in general. Nord University therefore encourages qualified candidates with disabilities, gaps in their CV, immigrant backgrounds or different life experience to apply. If applicants indicate they have disabilities or gaps in their CV, their details may be used anonymously for the purposes of governmental annual review Application process Applicants will be evaluated by an internal expert committee. Relevant applicants will be invited for an interview, trial lecture or other presentations. For more information, see Nord`s website . The Faculty reserve the right not to hire applicants who may be scientifically qualified but who do not meet other criteria such as language, educational background or personal characteristics Application Applications must be submitted electronically by 15.05.2026. The application should include: Application letter with description of your motivation for applying for the position Complete CV (education, work and teaching experience and overview of any scientific publications) Copy of the applicant’s master’s thesis Copy of diploma for both bachelor and master education and relevant certificates Applicants with education from countries outside Scandinavia with diplomas issued in a language other than English, must enclose witnessed diplomas in both the original language and authorized English translations Applicants from outside the EU/EEA must submit HK-dir approval of their master´s degree Project description of 3-5 pages where you explain the idea for the PhD project, including theoretical anchoring, methodological approach, expected outcomes and a clear time line of the whole PhD project. Name and contact information of 1-3 references (name, your relation to the person, email and phone number) Other activity that is considered relevant for the position, and wishes to be taken into account in the assessment All documents must be uploaded as attachments as a PDF files to the electronic application form. It is the applicant’s responsibility to submit the complete documentation within the deadline. Failure to include items mentioned above will automatically render the candidate unqualified. After the application period has expired a list of applicants will be published. Note that, in special cases, the identity of particular applicants to a position may be made public even when that applicant has requested that their application be processed confidentially. Applicants requesting an exemption from publication of their identity must explain their reasons for doing so in their application. In this case, advance notice will be given in the event of a decision to make any information about the applicant public. Apply Now

About the position Faculty of Biosciences and Aquaculture, Nord University, invites applications from outstanding candidates for a three-year PhD position in marine nature conservation and fisheries. The successful candidate will be part of the Ecology division, located in Bodø, Norway. This PhD will seek synergies between marine nature conservation and sustainable fisheries, notably by identifying opportunities for Marine Protected Areas to benefit fisheries through spillover benefits, as well as the MPA being refugia for broodstock and juveniles. Both global, regional and local perspectives and applications, across finfish and shellfish, are encouraged, with the aim to develop general theory around optimising marine spatial management to restore biodiversity while enabling sustainable use of resources. The preferred start date is September 1, but this is flexible. Qualification requirements A master’s degree in a relevant field of study is required Grade average of B or better (or equivalent score) both for the programme as a whole and the master’s thesis See admission requirements: PhD in Biosciences Employment is conditional upon admission to the faculty’s doctoral program. The final project outline must be approved within three months of appointment. Termination may be considered if admission to the doctoral program is not obtained within this deadline, or in cases of failure to submit annual progress reports and/or serious deficiencies in progress or duty work. For admission requirements and regulations, see our web page Fluent English skills are required both written and oral. A solid understanding of fisheries, biogeography, marine ecology, natural history, and nature conservation science and practice is essential. Strong computational and analytical skills, and ability to quickly learn new analytical methods are important. Experience in publishing papers as a lead author is an asset. Candidates who already have a PhD degree will not be considered for the position. Qualified applicants will be ranked according to the following criteria: Theme and perspectives from the applicant’s master’s study The applicant’s description of motivation for the doctoral work and opportunities for completion Relevant experience from research work Ability to work independently, goal-oriented and systematically Personal suitability We offer The annual salary for doctoral researchers (code 1017) is NOK 550 800,-. Benefits from the Government Pension Fund for loans, insurance and pension Creative and collegial working environment A workplace with progressive development Flexible working hours An active university sports club General information The person who is appointed must abide by the applicable laws, agreements, and directives. The responsibilities and duties associated with the position may change with future reorganization at Nord University. Nord University is subject to the Security Act, which sets requirements for the handling of classified and sensitive information. To increase security in the hiring process, Nord may conduct background checks on relevant candidates. Please note that all our candidates may be asked questions necessary in this context. This includes questions about any potential connection to countries that PST has identified as key threat actors in its annual threat assessment. The state workforce should, to the greatest extent possible, reflect the diversity of the population. Therefore, Nord University encourages qualified candidates with disabilities, gaps in their CV, immigrant backgrounds, and other diverse life experiences to apply. Information about disabilities or gaps in the CV may be used for anonymized registration purposes. For this position, Nord University is cautious about hiring individuals with connections to countries that PST has identified as key threat actors in its annual threat assessment. Contact information Further information about the position can be obtained by contacting: Mark Costello, mark.j.costello@nord.no Application process Applicants will be evaluated by an internal expert committee. Relevant applicants will be invited for an interview, trial lecture or other presentations. For more information, see Nord`s website . The Faculty reserves the right not to hire applicants who may be scientifically qualified but who do not meet other criteria such as language, educational background or personal characteristics Application Application deadline: 01.06.2026. The application should include: Application letter with description of your motivation for applying for the position Complete CV (education, work and teaching experience and overview of any scientific publications) Copy of diploma for both bachelor and master education and relevant certificates Applicants with education from countries outside Scandinavia with diplomas issued in a language other than English, must enclose officially certified diplomas in both the original language and authorized English translations Applicants from outside the EU/EEA must submit HK-dir approval of their master´s degree (Norwegian Directorate for Higher Education and Skills) Copy of the applicant’s master’s thesis Name and contact information of 1-3 references (name, your relation to the person, email and phone number) Other activity that is considered relevant for the position, and wishes to be taken into account in the assessment All documents must be uploaded as attachments as a PDF file to the electronic application form. It is the applicant’s responsibility to submit the complete documentation within the deadline. After the application period has expired a list of applicants will be published. Note that, in special cases, the identity of particular applicants to a position may be made public even when that applicant has requested that their application be processed confidentially. Applicants requesting an exemption from publication of their identity must explain their reasons for doing so in their application. In this case, advance notice will be given in the event of a decision to make any information about the applicant public. Apply Now

The Laboratory for Molecular and Translational Pharmacology (MolPharm) at Panum Institute, University of Copenhagen, is seeking a highly motivated PhD fellow to characterize viral membrane proteins. The PhD position is fully funded and for 3 years, starting on 15 July 2026, or as soon as possible thereafter. Information on MolPharm can be found at: Rosenkilde Group – University of Copenhagen Our research We investigate how membrane proteins translate structural features into functional outcomes. For many years, we have focused on G protein-coupled receptors (GPCRs) with an emphasis on ligand recognition, receptor activation, intracellular cycling, biased signaling, and crosstalk in the immune and endocrine systems. Our work extends to virus-encoded ion channels (viroporins), where we study the same properties as those of the receptors. We combine molecular pharmacology, structural biology, computation, and quantitative biophysics, supported by state-of-the-art biophysical assays and site‑directed approaches to identify and characterize novel modulators of GPCRs and viral ion channels. The project We are looking for a driven and ambitious person who can work independently and in collaboration with other groups. The candidate should have a strong interest in pursuing a PhD degree in the field of protein science with a focus on viral membrane proteins. The project will address fundamental questions regarding the structure, oligomerization, and pore-forming and/or ion‑channel function of viroporins, a diverse family of small viral membrane proteins that form oligomeric pores essential for viral life cycle and pathogenesis. The goal is to define the molecular principles governing viroporin assembly, map intra- and intersubunit contacts, and study pore function, ion selectivity, and druggability via modulation by small‑molecule inhibitors. This includes establishing high‑throughput pipelines for heterologous expression, purification, and reconstitution of viral membrane proteins. Druggability will be assessed using a high-throughput fluorescence-based screening platform in a human cell line. Who are we looking for? The person we are looking for should have a master’s degree or equivalent in biochemistry, structural biology, medicinal chemistry, molecular biomedicine, or a related field, with basic theoretical and practical knowledge of protein science, cell biology, and pharmacology. The applicant must be fluent in written and spoken English. The candidate is expected to take the initiative in driving the project while also collaborating with other members of the MolPharm lab. Emphasis will be given on personal qualities, including analytical capacity, accuracy, flexibility, and collaborative skills. Additional merits or desired skills: The ideal candidate has one or more of the following competencies: molecular pharmacological characterizations of membrane proteins in eukaryotic cells, including, for instance, GPCRs or ion channels; protein chemistry approaches such as recombinant protein expression and purification, biochemical and biophysical characterization of proteins; advanced microscopy methods. The selection is based on the following criteria: Completion of the above qualifications, grades in relevant courses, suitability of the courses to the research program, previous experience in working and/or performing research in the wider area that relates to structural and functional properties of membrane proteins. An integrated assessment of the applicant is conducted to identify the best possible candidate who can succeed in doctoral studies and contribute to the positive development of the department and the research program. Our group and research- and what do we offer? The Rosenkilde Group is part of the Department of Biomedical Sciences at the University of Copenhagen, located on the Panum campus in the Northern Campus area, close to the center of Copenhagen. We offer creative and stimulating working conditions in a dynamic, international research environment. The group forms an integral part of the Molecular and Translational Pharmacology research division, bringing together scientists with shared interests in membrane proteins: G protein‑coupled receptor (GPCR) biology, viroporins, ion channels, transporters, ligand recognition, receptor activation and desensitization, allosteric modulation, and oligomerization. Our research focuses on understanding the relationship between the structure and function of membrane proteins. We combine molecular pharmacology, structural biology, biophysical assays, and computational approaches to uncover mechanisms of activation, modulation, and inhibition, discover novel ligands, and refine them to improve drug properties. The environment provides access to state‑of‑the‑art experimental facilities for receptor characterization, ligand screening, and functional assays, supported by a broad expertise spanning protein engineering, mutagenesis, signaling analyses, and biophysical characterization Principal supervisor is Professor Mette M. Rosenkilde, rosenkilde@sund.ku.dk Job description: Your key tasks as a PhD fellow at SUND are: Carrying through an independent research project under supervision Completing PhD courses or other equivalent education corresponding to approximately 30 ECTS points Participating in an active research environment, including a stay at another research institution, preferably abroad Writing scientific papers aimed at high-impact journals Obtaining experience with teaching or other types of dissemination related to your PhD project Writing a PhD thesis on the grounds of your project Terms of employment The average weekly working hours are 37. The position is a fixed-term position limited to a period of 3 years. The employment is conditioned upon the applicant’s successful enrolment as a PhD student at the Graduate School at the Faculty of Health and Medical Sciences, University of Copenhagen. This requires submission and acceptance of an application for the specific project formulated by the applicant. The PhD study must be completed in accordance with The Ministerial Order on the PhD programme (2013) and the Faculty’s rules on achieving the degree. Salary, pension, and terms of employment are in accordance with the agreement between the Ministry of Taxation and the Danish Confederation of Professional Associations on Academics in the State. Depending on seniority, the monthly salary starts at approximately 31,200 DKK, /Roughly 4,170 EUR (November 2025 level). In addition, the employer will pay 18,07% in pension. The position is covered by the Protocol on Job Structure. Application procedure Your application must be submitted electronically by clicking ‘Apply now’ below. The application must include the following documents in PDF format: Motivated letter of application (max. one page) CV incl. education, experience, language skills, and other skills relevant for the position Certified copy of original Master of Science diploma and transcript of records in the original language, including an authorized English translation if issued in a language other than English or Danish. If not completed,